And here we get the 250 ml more PC volume. SInce the Mk1 bladder do expand more, there is more useable space in the PC casing. The Mk2 bladder expands not as much as the mk1, therefore you need more "base-bladder-lengh" to get the same PC, but the "base-lengh" needs space. And talking of the CPS 2000 Mk1 and mk2 this is the reason for the 250 ml PC volume difference. The MK1 bladder is even a bit SHORTER but expands MORE

But keep in mind that 250 ml is not much, so the difference is a short laugh

Furthermore we have to consider following thing: If we want a big stream with best range(50+) we will need two things :
Pressure and volume
The Mk1 bladder do contract faster than any other bladders, only this way we get a strong and steady stream with 50 + range. When we open the firing valve and we loose pressure, the bladder's contraction is there to keep the pressure stable. But the bigger the output/s the faster the bladder must expand to keep up the range. Speaking of the CPS 2000 ' nozzle we have a stream size where the mk2 bladder contracts just a bit too slow, other than the mk1, which expands just fast enough.
Therefore, using the BIG BALL VALVE of the HC, it is BETTER to use the mk1 bladder than the Mk2, since the mk1 contracts faster, which ends up in a great recoil and powerful stream

Hope you can follow me, not easy to transform one's thoughts into a text
